Well ... (remembering that I haven't used a Mac in 18 months so I might misremember details and things might have changed) ...
Anything you can get for UNIX is almost certainly going to be available (and the usual collection of FOSS stuff is free), but things that run in X11 don't tie in nicely with the rest of the system.
Apps designed for the Mac appear in the toolbar at the bottom of the screen, but anything running under X gets lumped into the icon for X. Drag and drop doesn't work smoothly between X apps and native apps. And so on...
This means that, generally, you'll be wanting to do things with native apps whenever possible - but most of these are not FOSS.
Then are the class of apps that fiddle with the GUI, so if you want (for instance) Focus Follows Mouse or Virtual Desktops, then you have to start forking out for software such as CodeTek Virtual Desktop.
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