$pi_sum = 0; $L_sum = 0; $differences_sum = 0; $coverage_sum = 0; for my $PAR1 (@PAR1) { my @PAR1_info = split(/\t/, $PAR1); $pi = $PAR1_info[0]; $pi_sum = $pi_sum + $pi; $L = $PAR1_info[1]; $L_sum = $L_sum + $L; $differences = $PAR1_info[2]; $differences_sum = $differences_sum + $differences; $coverage = $PAR1_info[3]; $coverage_sum = $coverage_sum + $coverage; my $PAR1_diversity = (($pi_sum/$L_sum)/($differences_sum/$coverage_sum)); }